Behold a result of uncontrolled greed and ambition.
Mainframe Prime is the only one of the Legendary Pokégirls that used to be human. She was born Linda McKenzie, and is the instigator that caused the downfall of James Scott with her misrepresentative reporting and his subsequent rise to infamy as Sukebe. She was captured by Macavity Prime, and brought before Sukebe to face punishment for her part in making him the man what is is today. She wasn’t discovered until after the end of the Titan War, however records were found that showed she had a part in it.
Mainframe, like all the Legendaries save Macavity, is a tall Pokégirl, ten feet in height. Her body is robotic, with a base similar to other robotic Pokégirls, with metallic skin and hair, and wires showing in her joint connectors, her breasts a large E-cup. Her shoulder armor resembles monitors, and her helmet is possessed of a wide visor that covers her eyes, data constantly flowing over it. Her chest and back armor is thick, mounted with over a dozen wires that allow her to uplink with any computer system on the planet and access EVERY computer on the planet. On the front are monitors showing constantly streaming lines of data. Her leg armor is equipped with jet thrusters on the boots, and her arm-mounted armor is thicker around the forearms, due to the energy emitters on them. In her chest, where her heart would be, is a power core of sorts that is more armored than the rest of her body. Even unarmored, her body is heavy despite her slender looks, making her overall movement speed slow. This doesn't work against her in deep water, as she can withstand great amounts of pressure, almost as much as a Deep Dive, and no longer needs to breath.
Linda McKenzie was loud, arrogant, and condescending. Mainframe Prime is quiet, contemplative, and thoughtful, a trait her daughters share. She prefers to be alone as much as possible, psychics noting a great sense of shame in her. She is frequently known to cry for no apparent reason, possibly due to what Sukebe did to her as punishment for her part in his fall. She is very aware of her part in the way the world was shaped, and while she is still bitter towards Sukebe, does not blame him. She possesses a rancid hatred of Macavity, though, one that Macavity shares after an incident caused my Mainframe in which the Cleric of Chaos was publicly humiliated and nearly killed. The two have had a bitter rivalry ever since, and are confused beyond measure by the fact that their daughters get along so well.
In addition to being turned into a sex-needing Pokégirl, Linda McKenzie was also put under a magical curse: she is aware of everyone that has died since the beginning of the Revenge War. When someone dies, she learns their name and their history, recording it in her internal database. It was the most fearsome punishment Sukebe could think of, to make sure that she remembered everyone who died 'because of her arrogance,' be they human or nameless Pokégirl that dies in the wild. She is known as 'She Who Remembers,' and in modern times is honored by some as one who will never let anyone be forgotten. Thankfully, for the sake of her daughters, the curse does not carry over to them. Over time, Mainframe has developed a hatred for those who desecrate graves and try to ruin the memories of those who died, a trait which has allowed her to form a friendship of sorts with fellow Legendary, Death. All the arrogance and greed which motivated Linda McKenzie has been stripped away, leaving only a humble Legendary who wishes nothing more than to gather data in peace.
Her daughters, DemiMainframes, do not have the same depression their mother does, however they are as quiet and introspective. They are known for doing what they can to make their harem sisters and Tamers happy, as their mother's perpetual sadness leaves them wanting to keep those they care about from crying. Like their mother, they have an infinite data storage capacity and flawless total recall, making them immune to amnesia effects and madness effects. Both Mainframe and her daughters, however, do have a tendency to develop behaviors consistent with OCD. This is usually manageable, however, and not disruptive to the overall harem dynamic.
Mainframe isn't the fastest of fighters, her heavy body hindering her in combat. But she can take a great deal of punishment due to her body's density. Her main method of attack is to use powerful strikes to decimate the opponent, along with powerful energy strikes. Her armor has unique energy emitters on the wrists, shoulders, hips, and cannons on her back, which can make copies of her data and transmute it into powerful energy attacks which take the forms of ones and zeros. She also possesses tentacles which she can use to uplink not just to computer systems, but to other Pokégirls, allowing her to learn their capabilities and think of a way to counteract them. The tentacles are even useful in combat and other areas.
Mainframe didn't like taming at first, according to journals found shortly after her discovery. When she was first transformed into a Pokégirl, Sukebe basically kept her as a pet, inflicting his memory curse on her and forcing her to watch as the Revenge War tore the world apart. When the Titans came, Sukebe put her to work, setting up the lab she was eventually found in and having her coordinate communications for both humans and Pokégirls, helping to rally them against the Titans and their monsters. She herself fought rarely, but the bones of several large breeds of Titan monster near her lair showed that she could fend them off when needed. After the Titan War and Sukebe's death, seeing no alternative, she decided to keep monitoring the world. She would only seek out Taming as a necessity, and at first wouldn't stay with a Tamer after being impregnated. But after the first of her daughters were born, she began to mellow out, her pride slowly melting away until she began to actually like having sex. She realized the benefits of her work, and began to learn about the living as well, gaining an appreciation for life as strong as her respect for the fallen.
Mainframe and her daughters are known for generally preferring more basic forms of sex. Missionary, anal, oral, things like that. Because of the density of their bodies, they generally prefer to be on the bottom. Her armor is removable, like all robotic Pokégirls, allowing for easier Taming. Mainframe and her daughters are also known for being very quiet in bed, not making much noise no matter how much pleasure they are in. Their breasts are known to be very sensitive and easy to stimulate, a good chest message a way of warming them up.
Mainframe generally kept to herself for the first few hundred years, even after she was officially discovered. She was rarely seen with a human, and kept her activities hidden. She slowly began to be seen outside her lab more often, and was even known to smile after the incident she arranged to humiliate Macavity. She dropped her more anti-social tendencies completely, however, after a failed attempt to commit suicide by fighting Menhit. The warrior Pokégirl defeated her easily, decimating her despite her superior armor and attacks, but refused to deal the deathblow. She then said something to Mainframe, which was not overheard. Whatever it was, it left Mainframe a sobbing wreck. Shortly afterwards, Mainframe began to take a more active role in her own life, journeying from location to location as her other sisters did, aiding them where necessary, and seemingly enjoying just existing more.
Shortly after her suicide attempt, a young Legendary Pokégirl was found in the water near the Chaos Zone. The Legendary Primes were the first to discover her, having sensed her arrival and immediately dropping whatever they were doing to track her down. It was the first recorded instance of all the Legendaries, including the vicious War, the stoic Death, and mad Macavity, showing fear, something that Macavity Prime later admitted hadn't happened since Typhonna turned on them. It was also the first time they had all worked together willingly since before Sukebe's death. After the Legendary Pokégirl, named Abyss, was studied and her origins discovered, Mainframe Prime was entrusted with her care. The tiny part of Mainframe Prime that still considers herself Linda McKenzie was amused by the irony of this...
It is not possible for someone to Threshold into a Legendary Pokégirl. Primes only become fertile once every five years, and DemiLegendaries only give birth to humans. Legendary Primes tend to be very protective of their daughters, raising them until they feel they are safe to leave her on their own. They are also constantly aware of their daughters, so anyone who catches a DemiLegendary can expect a guaranteed visit from the mother in short order. Somehow, they always know where the one who caught their daughter is, so hiding is impossible.
When a Legendary Pokégirl is ready to breed, they will attach themselves to the harem of a trainer that they think will be able to put up with them. They stay with him (and it will always be a male, although they are not above lesbian sex if it amuses their temporary master) throughout the duration of their pregnancy, listening to the orders of their temporary master until two months after they give birth. The gestation period of a Legendary Pokégirl is one full year.